eprintid: 1508144 rev_number: 29 eprint_status: archive userid: 608 dir: disk0/01/50/81/44 datestamp: 2016-11-14 15:34:38 lastmod: 2021-12-13 02:18:55 status_changed: 2016-11-14 15:34:38 type: article metadata_visibility: show creators_name: Hannan, S creators_name: Gerrow, K creators_name: Triller, A creators_name: Smart, TG title: Phospho-dependent Accumulation of GABABRs at Presynaptic Terminals after NMDAR Activation ispublished: pub divisions: UCL divisions: B02 divisions: C08 divisions: D09 divisions: G02 keywords: Science & Technology, Life Sciences & Biomedicine, Cell Biology, GABA(B) RECEPTOR INTERNALIZATION, GAMMA-AMINOBUTYRIC-ACID, CENTRAL-NERVOUS-SYSTEM, PROTEIN-KINASE, LATERAL MOBILITY, INHIBITORY NEUROTRANSMISSION, HETEROSYNAPTIC DEPRESSION, SURFACE TRAFFICKING, SYNAPTIC PLASTICITY, SUSHI DOMAINS; AMPK; bungarotoxin binding site; Ca2+ signaling; excitotoxicity; GABA receptors; GABAB receptors; glutamate receptors; hippocampus; homeostatic signaling; lateral diffusion; NMDA receptors; phosphorylation; presynaptic terminal; quantum dots; receptor mobility; single-particle tracking; sushi domains note: © 2016 The Author(s). This is an open access article under the Creative Commons Attribution 4.0 International (CC BY 4.0) license (http://creativecommons.org/licenses/by/4.0/) abstract: Here, we uncover a mechanism for regulating the number of active presynaptic GABAB receptors (GABABRs) at nerve terminals, an important determinant of neurotransmitter release. We find that GABABRs gain access to axon terminals by lateral diffusion in the membrane. Their relative accumulation is dependent upon agonist activation and the presence of the two distinct sushi domains that are found only in alternatively spliced GABABR1a subunits. Following brief activation of NMDA receptors (NMDARs) using glutamate, GABABR diffusion is reduced, causing accumulation at presynaptic terminals in a Ca2+-dependent manner that involves phosphorylation of GABABR2 subunits at Ser783. This signaling cascade indicates how synaptically released glutamate can initiate, via a feedback mechanism, increased levels of presynaptic GABABRs that limit further glutamate release and excitotoxicity. date: 2016-08-16 date_type: published publisher: CELL PRESS official_url: http://doi.org/10.1016/j.celrep.2016.07.021 oa_status: green full_text_type: pub language: eng primo: open primo_central: open_green article_type_text: Article verified: verified_manual elements_id: 1148658 doi: 10.1016/j.celrep.2016.07.021 lyricists_name: Hannan, Saad lyricists_name: Smart, Trevor lyricists_id: SBHAN72 lyricists_id: TSMAR12 actors_name: Smart, Trevor actors_id: TSMAR12 actors_role: owner full_text_status: public publication: Cell Reports volume: 16 number: 7 pagerange: 1962-1973 pages: 12 issn: 2211-1247 citation: Hannan, S; Gerrow, K; Triller, A; Smart, TG; (2016) Phospho-dependent Accumulation of GABABRs at Presynaptic Terminals after NMDAR Activation. Cell Reports , 16 (7) pp. 1962-1973. 10.1016/j.celrep.2016.07.021 <https://doi.org/10.1016/j.celrep.2016.07.021>. Green open access document_url: https://discovery.ucl.ac.uk/id/eprint/1508144/1/smart_1-s2.0-S2211124716309202-main.pdf