%D 2023 %O © 2023 Zhang et al. This work is licensed under the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/). %V 133 %I American Society for Clinical Investigation %L discovery10174707 %C United States %N 12 %T KIR-HLA interactions extend human CD8+ T cell lifespan in vivo %A Y Zhang %A AWC Yan %A L Boelen %A L Hadcocks %A A Salam %A DP Gispert %A L Spanos %A LM Bitria %A N Nemat-Gorgani %A JA Traherne %A C Roberts %A D Koftori %A GP Taylor %A D Forton %A PJ Norman %A SGE Marsh %A R Busch %A DC Macallan %A B Asquith %J Journal of Clinical Investigation %X BACKGROUND. There is increasing evidence, in transgenic mice and in vitro, that inhibitory killer cell immunoglobulin-like receptors (iKIRs) can modulate T cell responses. Furthermore, we have previously shown that iKIRs are an important determinant of T cell–mediated control of chronic viral infection and that these results are consistent with an increase in the CD8+ T cell lifespan due to iKIR-ligand interactions. Here, we tested this prediction and investigated whether iKIRs affect T cell lifespan in humans in vivo. METHODS. We used stable isotope labeling with deuterated water to quantify memory CD8+ T cell survival in healthy individuals and patients with chronic viral infections. RESULTS. We showed that an individual’s iKIR-ligand genotype was a significant determinant of CD8+ T cell lifespan: in individuals with 2 iKIR-ligand gene pairs, memory CD8+ T cells survived, on average, for 125 days; in individuals with 4 iKIR-ligand gene pairs, the memory CD8+ T cell lifespan doubled to 250 days. Additionally, we showed that this survival advantage was independent of iKIR expression by the T cell of interest and, further, that the iKIR-ligand genotype altered the CD8+ and CD4+ T cell immune aging phenotype. CONCLUSIONS. Together, these data reveal an unexpectedly large effect of iKIR genotype on T cell survival.